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When Doctor Thorne's penniless niece, Mary, is excluded from the wedding preparations of her childhood playmate, she probes her uncle for the truth of her birth. She is devastated to learn that she is the illegitimate child of his late brother. It seems she has neither breeding nor fortune. This is a problem, as Mary and the heir to the Greshamsbury Estate, Frank, have fallen in love. Frank is under strict instructions from his over-bearing mother, Lady Arabella, to save his family from financial ruin by marrying money. Frank's mother schemes to separate the two lovebirds, encouraging her son to woo the wealthy American heiress, Miss Dunstable.